Brian,
I do think the calf/ankle complex is likely better suited to take the force of running by design.
Note the lack of ankle replacements these days (like they even could), compared to the excess of knee replacements. I hardly see any ankle arthritis, while I see a ton of knee arthritis. I'd say that knees are wearing out prematurely compared to ankles, possibly showing that the knees are being "used" incorrectly...
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