~4000-4300/day according to fitday, "Just north of vag" variant of 531, I ran once a week and NEPA included bouldering 2x. I do "throw off heat", but again, my theory was insulin management was so tight I simply wasn't storing/using much of the calories.
I've done a couple of 20RM squat exercises.. the after effects are quite interesting.
The point I forgot to make about 531 and the volume was that I am far too much of an amateur for it to be the best program. I had (and have) a ton of weight/strength to gain before a true intermediate program becomes the best available option.
Anyway, just my experience/2 cents.